Galaxy source includes the Galaxy interfaces but not the third party executables for tools like fastqc or bwa. They can be automagically installed if you install the tool from a tool shed but at a guess, you are working on your desktop with the fastqc tool in a recent clone of galaxy? Unfortunately, the tool can't run until you have the fastqc software working in a particular way hinted at in the guide for the fastqc tool on the page at
http://wiki.galaxyproject.org/Admin/Tools/Tool%20Dependencies - try that please?